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%SystemRoot%

A typical value for the environment variable %SystemRoot% seems to be C:\Windows. %windir% is another environment variable that also points to this location.
Among others(?), this (and its subdirectories?) store so-called System apps which are integral to the Windows Operating System. See also types of apps in Windows 10.
%SystemRoot% is also found in the registry as value of Directory under the registry key HKEY_LOCAL_MACHINE\SYSTEM\CurrentControlSet\Control\Windows.
The value of %SystemRoot% can be queried with
wmic os get WindowsDirectory

Opening folders below %SYTEMROOT%

Folder below %SYTEMROOT% can be opend in cmd.exe with start (if %SystemRoot% is in %PATH% which it usually is). The following example opens the fonts folder below C:\windows
C:\> start fonts
Similarly, these folders can also be directly opened using Win+R and then typing the folder name.

Relationship between %SystemRoot% and %windir%

%windir% is supposed to the same directory as %SystemRoot%.
The difference between these is that %SystemRoot% is not a real environment variable but is kind of hardcoded into the kernel while %windir% is an ordinary whose value is set in the registry under HKEY_LOCAL_MACHINE\SYSTEM\CurrentControlSet\Control\Session Manager\Environment and usually assigned the value of %SystemRoot%.
The value of %SystemRoot% is determined when Windows is installed.
Allegedly, in Windows 10, the environment variable %CSIDL_WINDOWS% also points to %SystemRoot%. However, I found that not to be the case.

Some files

bfsvc.exe Boot File Servicing Utility
bootstat.dat
DtcInstall.log
explorer.exe Windows Explorer
HelpPane.exe Microsoft Help and Support
hh.exe Microsoft® HTML Help Executable
lsasetup.log
mib.bin
notepad.exe Notepad
PFRO.log
Professional.xml
regedit.exe GUI based registry editor.
setuperr.log
splwow64.exe Print driver host for applications
system.ini
twain_32.dll Twain_32 Source Manager (Image Acquisition Interface)
win.ini
WindowsUpdate.log
winhlp32.exe Windows Winhlp32 Stub
WMSysPr9.prx
write.exe Windows Write
Some subdirectories of %windir% might be:

Showing the «System Root» with bcdedit

The value of the System Root shows up, for example, when displaying boot options with bcdedit.exe:
C:\> bcdedit.exe /enum
…
Windows Boot Loader
-------------------
…
osdevice                partition=C:
systemroot              \WINDOWS

TODO

Compare with the values PathName and SystemRoot under KEY_LOCAL_MACHINE\SOFTWARE\Microsoft\Windows NT\CurrentVersion.

See also

The .NET enumeration System.Environment+SpecialFolder.
%SYTEMROOT%\INF
The value of Windows Directory under System Summary in msinfo32.exe
get-computerInfo | select-object osWindowsDirectory

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